Kadua cookiana

Cham. & Schltdl. (1829)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Rubiaceae Genus: Kadua

‘āwiwi [awiwi]

Description

Key Characters:

Growth Form: Small, many-branched shrubs.

Stems: Stems striate, to ca. 1–2 dm long, glabrous.

Roots:

Leaves: Leaves simple. Opposite. Blades linear, 4–8 cm long, ca. 0.5–1.2 cm wide. Blades chartaceous. Margins slightly revolute. Veins other than midrib obscure. petioles 0 – 0.5 cm long, connate into a sheath. Stipules reduced to a carinate, linear–subulate mucro 2–5 mm long, adnate to petiole bases.

Flowers: Flowers 3(–7) in cymes, these hirtellous throughout or glabrous, pedicels ca. 8–15 mm long, those of the central flower longest, bracts subtending inflorescence linear, ca. 1–1.5 cm long; hypanthium turbinate, ca. 1.5 mm long. Flowers insect-pollinated, apparently bisexual (perfect) or pistillate. Calyx 4–5 lobed, lobes linear–deltate, 4–6 mm long, 1.5–2 mm wide, apparently not enlarging in fruit. Corolla white, fleshy, salverform, the tube ca. 8–9 mm long, the limb not strongly quadrangular in bud, but apex depressed; 4-5-lobed, the lobes ca. 2 mm long at anthesis, strongly inflexed in bud; nectary disc present. Stamens as many as and alternate with the corolla lobes, inserted on corolla tube midway or near throat; anthers sessile or on short filaments; anthers dithecal, opening by longitudinal slits. Ovary inferior or partly so, 2(-4)-locular; ovules few to numerous on fleshy placentas near middle of septum; styles as many as carpels, cylindrical to filiform, terminal, slender; stigma bilobed or subcapitate, included or exserted.

Fruit: Capsules turbinate–globose; 3–3.5 mm long; 3.5–4 mm in diameter; dehiscing loculicidally across disk; tardily slightly further; pyrenes tardily dehiscing septicidally; endocarp thinly sclerified; disk conical to slightly beaked. Seeds reddish brown; cuneiform; slightly compressed; irregularly angled; with well-developed oily endosperm; or endosperm occasionally scanty or absent.

Ploidy:

Habitat: Probably in lowland habitats.

Elevation Range:

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
Hawai'i Extirpated

Dispersal Agents


Pollinators

Bibliography

Name Published In: Linnaea 4: 158 (1829)

Other References

Wagner et al. 1990:1140 (K, H [as Hedyotis cookiana]); Terrell et al. 2005:833/Kennedy et al. 2010:23 (Syn. H. cookiana = K. cookiana); Chamisso s.n. B (H†, 1817)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Plants 6-15 inches long. Flowers white. Christensen, C. 6 Kauai BISH 1976-05-08
2 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details mixed mostly alien vegetation with Bidens forbesii, Carex wahuensis, Psidium guajava, Erigeron karvinskianus, Ageratum conyzoids, Blechnum occidentale, Clidemia hirta, Lantana camara, Lythrum maritimum, Kalachoe pinnata, Melinus Preserved_Specimen 17 adult plants, about 50 juvenile plants; seedlings 1 ft ht with white flowers Perlman, S. 21690 Kauai BISH 2009-06-15
3 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details riparian vegetation, with Kadua cookiana, K. elatior, Poa mannii, Plantago princeps longibracteata, Deparia cataracticola, Isachne pallens, Asplenium unilaterale, Lipochaeta connata, Selaginella arbuscula, Eragrostis variab Preserved_Specimen Sub-shrub, few branched, in flower, corolla white, ca. 50 plants Wood, K.R. 16499 Kauai BISH 2015-06-24
4 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Face of waterfall. Preserved_Specimen Flower white. St.John, H. 24966 Kauai BISH 1953-01-03
5 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Moist crevice at the base of the falls. Preserved_Specimen Flowers are white. St.John, H. 23143 Kauai BISH 1948-01-01
6 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Moist crevice at the base of the falls. Preserved_Specimen Flowers are white. Britten, E.J. 23143 Kauai BISH 1948-01-01
7 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen pendent stem, leaves chartaceous; calyx green, corolla white; style included Lorence, D.H. 8817 Kauai BISH 2001-11-09
8 Kadua cookiana Locality redacted. Contact Bishop Museum Botany Department for details Preserved_Specimen Prennial herb with decumbent, trailing stems, woody at base. Lvs. Plain dark green. Corolla pure white, no fragrance noted at any time. No fruit set observed. Lorence, D.H. 10173 Kauai BISH 2010-03-18